Pacheco, Thaynara L., y Fernando Z. Vaz-de-Mello. «Dung Beetles of the Tribe Phanaeini (Coleoptera: Scarabaeidade: Scarabaeinae) from Roraima State, Northern Brazil: Checklist and Key to Species». Biota Neotropica 15, no. 2 (junio 1, 2015). Accedido abril 26, 2025. //www.biotaneotropica.org.br/BN/article/view/1302.