Ferreira, K. R., and S. R. Rodrigues. “The Mating Behavior of Leucothyreus Marginaticollis Blanchard, 1843 (Coleoptera: Scarabaeidae: Rutelinae)”. Biota Neotropica, vol. 17, no. 4, Jan. 2017, //www.biotaneotropica.org.br/BN/article/view/1478.